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Soil structure is concerned with the living space in
soil; the size, shape and presence of the pore space. Similarly, soil
texture is concerned with the nature and working properties of the soil
particles, which by the manner of their arrangement enclose the living
space. The term "soil structure" and "soil texture"
are frequently confused. When assessing the quality of cloth by its
feel, we should try to distinguish between characteristics determined by
the nature and thickness of the thread and characteristics determined by
the nature and closeness of the weave. Similarly, when feeling soil, we
should try to differentiate between characteristics determined by the
nature and size of the fundamental particles by the manner of their
arrangement. In soil, texture is primarily a matter of particles;
structure and their arrangement. Texture is a fundamental, stable
characteristic of a soil that the farmer has to accept; structure is
temporary and can be manipulated. In a fertile soil the living space
consists of an abundance of small pores retentive of moisture and an
interconnected system of large pores that drain to admit air. The ideal
soil would have about 30% of its pore space unblocked by water and well
aerated. The rest of the pore space (70%) would be full of water
available to plant roots. The problem is to achieve the ideal mozaic of
air-filled and water-filled pore space in a medium consisting of
particles all roughly spherical, blocky or platy in form. When soil is
compact, particle size controls pore size, and hence, the rate of water
flow and the tenacity with which water will be held.
In soil structure formation, large particles
generally provide large pores in the soil system and thus permitting a
rapid flow of excessive moisture through the soil, i.e. rapid drainage,
but this advantage in respect of drainage is cancelled by the inability
of large pores to retain moisture against the pull of gravity once the
excess moisture has been cleared off the surface. In a large particle
system, therefore, there is a risk of an inadequate moisture reserve to
sustain plant life over periods of drought. Small particles provide
small pores resulting in much slower rates of water movement and a much
greater tendency for the pores to remain water-filled, once excess
moisture has been cleared through them. Therefore, a fine particle soil
will drain slowly and the pore system may well remain completely
waterlogged after drainage has ceased unless, by some means, large
diameter, free-drainage channels can be established through it. Organic
materials increases the total porosity with a decrease in bulk density
increases cation exchange capacity (CEC).
Decomposition of different organic amendments may be
affected by organic matter characteristics in terms of soil properties
such as soil texture, structure and pH. Soil structure influences soil
moisture, air-water ratio, capacity to infiltrate, conduct store and
drain water in addition to plant root growth development and
penetrations. Soil is formed from the weathered products of rocks
resorted and redistributed by the forces of gravity, wind, water and
ice. The end product is a surface mantle of loose mineral debris
consisting of particles far from uniform in shape and size. Neglecting
the very large particles that we might all agree to call either
boulders, stones or gravel there is, within what soil scientists define
as the "fine earth", a thousand-fold range in particle size.
The largest particles included within the sand category are just under 2
mm. in diameter (2000 micron). The smallest particles within the range
of particle size, which includes the clay minerals, are 0.002 mm (2
micron) in diameter. A useful and generally accepted system of defining
soil particle size categories is given below: - 2000-1000 mm, coarse
sand; 1000-500 mm, coarse sand; 500-250 mm, medium sand; 250-125 mm,
very fine sand; 125-50 mm, very fine sand; 50-20 mm, coarse silt; 20-2
mm, fine silt; less than 2 clay.
All the sands feel gritty and make a rasping sound
when rubbed between the fingers. The sharp, searing sound of the word
"sand" reflects this property. The silts, by contrast, feel
smooth and powdery when dry, oily smooth and slimy when wet. Again the
material being described seems to have the characteristics implied by
the sound of the word evolved over time to identify it. The sands and
the silts are essentially the product of the physical commination of
rocks. They are particles all roughly spherical or cubical in shape
hence, the finest of them, the very fine sands and the silts, readily
flow when saturated, clogging pores between larger particles, in-filling
drains and lubricating the erosion of soil down-slope. The clay minerals
are the particles typical of the finest particle size category
recognized in a textural description of soil, but clay particles are not
just very small particles less than 2 microns in diameter, and they are
not simply very fine particles, the ultimate product of physical
commination. Clays are formed directly from rock minerals by chemical
transformation, and considerably in form, chemical composition and
physical properties according to their mineral origin and the conditions
under which they were formed. Like the rust on iron they are not just
small particles of iron but are new materials formed by chemical changes
involving the regrouping of atoms.
However, the change from rock mineral to clay mineral
is not simply a change in particle size. This change also involves a
change in particle shape and in some important physical properties of
considerable significance for the development of structure. Clay
minerals generally are platy in form and like sheets of glass, cohere
when stacked moist whereas silt particles, which are more isodiametric
in form, would slump like a pile of minute billiard balls. But clay
particles not only stick to each other; they can serve as a binding
agent between other particles. Thus the clay fraction, if present as at
least ten percent of the whole, can confer to the soil, when moist,
qualities of stickiness and stiffness, and a degree of rigid cemented
when dry.
But a stack of platy clay particles has other
properties than cohesiveness: it will swell when able to absorb water
freely and shrink again as it dries out. On drying, clays will tend to
cause shrinkage to the point of cracking and the creation of large
compact clods but on re-wetting, moisture will cause swelling and clods
will tend to disintegrate into fragments producing a tilth similar to
that achieved by frost. With clay contents of thirty percent and more,
therefore, soils will, to some extent be self- mulching. Ultimately, of
course, excess moisture can cause such a tilth to collapse completely
but providing drainage is adequate to allow rapid clearance of excess
moisture from the large cracks that separate the clods, this is unlikely
to occur. Finally, clay particles, unlike sand and silt particles, are
charged. Like little magnets they have another mechanism by which they
can form links with other particles, and this mechanism, though not
rigid, can still function through water. But the charge is negative and
since like charges repel, this would seem to favour the dispersion of
clay particles rather than their clumping. However, there are many
positively charged particles in soil, in particular, calcium ions that
dissociate from the calcium carbonate in lime, and these by being
attracted to the negatively charged surfaces of clay particles, can
serve as a bridge between them, causing them to clump into loose
aggregates large enough to settle out of fluid suspension, like curds in
milk. Thus, though clay particles, were they to function as individual
units, would be small enough clay particles, were they to function as
individual, would be small enough to be washed out of the surface of the
soil and form drainage pans in the subsoil, this is unlikely to happen,
unless the lime status of the soil is allowed to fall.
The clay fraction, therefore, is a very important
fraction; not indispensable, as we shall see later, but very important
where it is present in sufficient quantity to influence working
properties. Soils with over forty percent clay are likely to be sticky
when wet and hard when dry, but also, they may be self mulching and
easily cultivated to a satisfactory, cracking tilth if worked at the
right moisture state. Given good subsoil drainage plant roots can
penetrate the large, well-aerated cracks, keeping them open and
deepening the zone of biological exploitation. Clay soils are naturally
retentive of moisture and of nutrients, and both can be exploited for
crop production provided weather conditions and cultivation techniques
preserve and encourage the cracking structure, required for drainage,
aeration and deep root penetration.
The sands and the silts provide only a mechanical
framework which, were it not for the cohesive properties of clay, would
be liable to slump into a system of particles individually packed to the
densest possible arrangement. If the predominant particles are of sand
size the soils, though quick draining and loose enough to be easily
worked will be thirsty and hungry. With insuff'cient small pores they
will be unable to hold an adequate reserve of water and, without
negatively charged surfaces on which to hold positively charged, basic,
nutrient ions such as calcium, potassium and ammonia, will rapidly
frequent coddling with carefully metered doses of water and nutrients.
Alternatively, they need reinforcing with organic matter.
Soils strongly biased in texture towards sand, silt
or clay are not typical of our soil. There are many soils with less than
seventy per cent sand and less than forty per cent clay. These, as shown
on the triangle of texture are the loam. Typically they contain twenty
to sixty percent silt. For example, the loam, clay loam and silty clay
loam of central Wales contain up-or-down of 35 %, 25 % clay and 40 %
silt. In the loam, the nature of the living space is greatly affected by
the possibilities that exist for inter-particle packing and the
uniformity with which this is achieved. The clay content is insufficient
to confer much stability to any favourable clustering of particles that
could be expected to occur in a heterogeneous mix so, unless
periodically disturbed by cultivation, they are liable to pack into a
close packing arrangement. In this state they will have fine silt
particles blocking any large pores created by the close packing of the
sand and will behave, in their pore characteristics, as if they were
dominantly silts. In fact, under these conditions, the sand particles
present would appear to be the equivalent of inert filler, occupying
space but not contributing to drainage or water retention. This no doubt
accounts for the fact that some of our soils that are almost entirely
silt are not as sow to drain as might be expected. It may also explain
why some of our problem sports filed soils are structurally degraded,
close packing, sandy loam.
The loams are like the sands in their readiness to
slump into close packing arrangements but, unlike the sands, and like
the clays, when close packed, they are dominated in their moisture
characteristics by particles so fine that drainage cannot be other than
slow and water is retained, after drainage has ceased, to an extent
detrimental to aeration. These, then, are the soils that would appear to
be most at risk to the hazards of structural collapse. These are the
soils to which British agriculture as a whole has had to become adjusted
and it would be surprising, therefore, if our traditional methods of
farming these soils not cherished accordingly. So far I have only
considered the conclusions that seem to arise from what we know of the
properties of the mineral skeleton, but most soils consist of more than
mineral particles. A pile of mineral particles is not a soil until it
has acquired a biological component of living and decaying organic
matter. Nitrogen is not a component of the normal range of rock minerals
and is built into the soil largely as a result of the activities of soil
organisms. For this important element at least, therefore, natural soils
are dependent on the presence of the organic fraction. However, it may
be we need not do things Nature's way and, with the advent of chemical
fertilizers, it could be argued that the organic fraction is no longer
essential for the supply of nutrients. Likewise, though organic matter
would seem to be of benefit to sands for water and nutrient retention it
could be argued that well-managed, clay soils, such as those at
Rothamsted, for example, are not dependent on organic matter either for
nutrients or water. However, for the loam, though organic matter is not
essential for water or nutrient retention, it is absolutely essential
for structure.
The unstable but open structure so essential for the
aeration of loamy soils is a consequence of particle aggregation —
granulation, if you like, rather than disintegration. Unlike the purely
physical process of clod disintegration by cracking, granulation is a
biological process involving organisms and ephemeral organic agents,
which arise as, intermediate products in the process of organic decay.
The effect is summarized diagrammatically. Thus a small particle system
that would form a uniform network of small, water-holding pores in close
packing is converted into aggregates of sand or gravel size with small
pores retentive of moisture within the aggregates and large pores for
drainage, aeration and root penetration between. The organic bonding
agent involved in this type of aggregation is not very strong but, if
the aggregate is not physically sheared, it remains effective in water.
The bonding is electrostatic involving negatively charged sites on long
chain organic molecules capable of threading together clay particles
through the intervention of positively charged bridging particles such
as calcium ions. However, the effect of the organic bonding agents is
only temporary, as they are themselves subject to microbial attack. They
need to be periodically replaced and for this, the soil needs to be
continuously replenished with fresh organic residues. These residues
must then be decomposed but not too rapidly or, as in the Tropics, their
effective life will be too short to be of any consequence for structure.
In addition, fresh organic residues must not only be decomposed to
provide bonding materials, to be effective, these materials must be
mixed into the soil by some churning agent. Being gums, they are
unlikely to be able to migrate into position of their own accord.
The most effective natural, churning agent for soil
is the burrowing earthworm. These organisms are present to the extent of
some fifty to a hundred per square yard in a fertile soil and may
process each year through their gut some twenty tons per acre of soil.
The end product of their activities is not only their numerous deep
burrows but soil, plus calcium from a special lime gland in their gut,
intimately blended with fresh organic residues and moulded into a
continuous cylindrical ribbon that coils into a water-stable, open cast
easily permeated by roots. These casts, though obvious when they appear
above ground, ate also found below ground and, when reinforced and
maintained by root binding, are responsible for the crumb-sized,
aggregate tilth that is a feature of well-managed, permanent pasture.
Cracking, root penetration and careful cultivation no doubt can all play
their part in preparing the way for earthworm activity and reinforcing
its effects but, for the job of positively creating the water-stable
aggregation so vital for loams, the earthworm is essential.
Worms are very sensitive to acidity and are absent
from soils with a pH of less than five. They are also unable to cope
with long immersion in water. As food they seem to prefer the fresh,
soft tissues of leaves. If they are to be encouraged, therefore the soil
needs first to be drained to avoid persistent water logging then brought
to an adequate lime status and cropped in a manner ensuring an adequate
return to the soil of fresh, or only partially decayed, organic
residues. To summarize: for healthy, vigorous plant growth we should aim
to create soil conditions favourable to the thorough exploitation of the
soil in depth, both for its moisture and its nutrient reserves. This
seems to require an actively growing root system that can forage for
itself. To achieve the desired root growth the pore space system must be
such as to allow for easy root penetration through a network of
relatively large, well-aerated pores within a structure of fine
particles that are water-retentive. A compact, sandy soil might
approximate to this when heterogeneously packed, and so might a strong
clay when split open by intensive cracking, but the loams, unless
aggregated, have too little sand to avoid being completely blocked by
the fine material present and too little clay to form an adequately fine
cracking structure. For the loams, therefore, Nature has had to evolve a
system of fine particle clustering and it is this that is achieved by
organic bonding agents organized through the casting activities of
earthworms. For the farmer this means he must try to maintain conditions
favourable to earthworms and then plan his management so as to preserve
their handwork, avoiding excessive foot vehicle traffic over the surface
when the soil is wet, and using skill in the timing of his cultivations.
If a farmer can maintain his soil both adequately aerated and adequately
watered in depth, he can then enjoy the consequences of deep, exploitive
rooting in the robust growth of healthy plants.
THE GENESIS OF SOIL STRUCTURE
Stable aggregation requires that individual soil
particles do not disperse in water, but aggregation is a step beyond
this to the creation of peds, which exist as units within the soil mass,
i.e. the creation of aggregates out of flocculated particles.
CEMENTATION PROCESSES
The Influence of Lime:
Lime, organic matter an adequate supply of fine textured particles are
generally considered to be the essentials for aggregate formation: i)
Ca++ will ensure clay flocculation but it is not essential for this
purpose as H+ is as effective and even Na+, in the presence of an excess
of NaC1 in solution, will not achieve dispersion; ii) Calcium humates,
though stable, are no more stable than H- humates; iii) Calcium seems to
assist in stable aggregation by acting as a cation bridge between
negatively charged sites on clay particles and similarly charged sites
on organic matter. However, it has been reported that hydrogen saturated
humus forms more stable aggregates with clay than calcium saturated
humus; iv) Little mention is ever made of the possible role of
earthworms in aggregated formation but if a mixing agent is essential
then it is not without significance, that an adequate level of calcium
is required for the presence of burrowing earthworms; v) Calcium,
through its influence on pH and the churning activity of earthworms can
have an influence on the decomposition of organic matter and therefore,
perhaps, the production of organic compounds with an influence on
aggregation.
The
Influence of Clay: Clay
particles, being plate-like, will become oriented parallel to each other
and to other flat surfaces on drying. When dry it forms a very effective
cement. Rehydration requires considerable agitation. A degree of
cementation is apparent long before the air dry state has been reached.
Oriented water molecules may be involved in linking clay particles
through bridging cations. The tenacity of such links will increase with
dehydration and finally direct links may develop between adjacent clay
surface through peripheral hydroxyls or bridging cations. The smaller
the clay particles and the smaller the adsorbed cations the harder the
aggregates.
THE INFLUENCE OF ORGANIC MATTER
i)
From a detailed study of the clay and organic matter contents of
water-stable aggregates it has been reported that organic colloids not
only cause a high degree of aggregation of clay particles but also
produce large aggregates; ii)
Only in the wet tropics does organic matter seem to be an insignificant
factor in stable aggregate formation. There dehydrated oxides of iron
and aluminium act as the principle cements; iii)
Hydrogen saturated colloidal organic matter will form effective stable
aggregates with clay, fine quartz or orthoclase, the link being a
consequence of electro-station forces, stabilized by dehydration. The
aggregation of calcium saturated organic matter is slightly less stable;
iv)
Direct link may be possible between negatively charged organic molecules
and positively spots along the broken edges of clay particles; v)
It has been suggested that the final stages of dehydration in base-rich
soils involving the migration of alkali soluble organic colloids to the
drying surface of the aggregate which on dehydration is to confer a
measure of stability. Under acid conditions the end-product of a drying
phase might be a surface coating of iron and aluminium oxides; vi)
Fresh or only partly decayed organic matter is much more effective in
promoting stable aggregation than stable soil humus. Thus, if
granulating substances are involved they would appear to be ephemeral
products of decay.
THE INFLUENCE OF MICROORGANISMS
i)
Soil bacteria will remain attached to soil particles or dispersed in the
soil solution according to the pH of the medium. Under acid conditions
they may be leached from the soil. Thus, the proteinaceous bodies of
soil bacteria may themselves act as a cementing agent when the medium is
not acid; ii)
Fungal hyphae, like fine roots, may contribute an element of mechanical
binding particularly effective in the stabilization of large aggregates;
iii)
As already discussed, with reference to the influence of calcium, many
products of organic decay may contribute to aggregation through ionic
linkages to charged mineral particles, possibly involving bridging
cations of oriented water molecules; iv)
Polysaccharide gums, apparently synthesized by micro-organisms when
stimulated by the addition of sucrose, have been shown experimentally to
have a marked effect on the stability of aggregates (i.e. when mixed
with soil and moulded into aggregates). Such organic compounds would be
readily attacked by other soil organisms and the duration of their
effect would depend, therefore, on the general level of microbial
activity. Where this is high throughout the year, as in the moist
tropics, polysaccharide gums are likely to survive long enough to have
any great influence on aggregate stability. Under the extreme
continental conditions of Steppe and Prairie countries there may well be
two periods of reduce microbial activity — one due to summer drought
and another due to winter cold. Under these conditions an adequate
reserve of stabilizing polysaccharide gums may be preserved and may
account for the very stable structure of natural, organic rich,
chernozem soils.
The Influence of Iron and Aluminium Colloida:
Ferric hydroxide precipitated as an hydrated gel at pH's around neutral
will become an effective, stable cement by dehydration. As dehydration
will take place at the periphery of fine textured structural units this
process will tend to create a stable skin around such aggregates but in
coarse textured materials, the iron will be concentrated in residual
moisture films around particles and pore outlets. These contrasting
end-products are to be seen in laterites and the B horizons of podzols.
Colloidal alumina probably behave in a similar manner.
The Influence of the Type of Clay Present:
Kaolinite, in the
absence of significant amounts of montmorillonite, hUmus, calcium or
iron oxides will, with alternate wetting and drying, tend to produce a
platy structure. This no doubt accounts for the platiness of the A2
horizon in podzols. All the above processes have really been concerned
with cementation after aggregate formation has taken place. The
processes now to be considered are those likely to contribute to
fragmentation or granulation.
FRAGMENTATION PROCESSES
The Influence of Alternate Wetting and Drying
i)
Drying of a moist soil mass will cause shrinkage, which if it is uneven,
will cause cracking. At a later stage dehydration of soil colloids will
result in cementation within the separated fragments; ii)
Re- wetting involves the dragging of water through fine pores and
between platy sheets by capillary forces with consequent swelling and
compression of occluded air; iii)
The bouyancy of occluded air will contribute to aggregate disintegration
should the method of re-wetting be rapid immersion in water. Wetting by
capillarity from below may well allow time for occluded air to be
released through the existing system of large pore spaces; iv)
Infiltrating with a non-polar liquid such as carbon tetrachloride will
reduce the disruptive effect of wetting by reducing the affinity of the
infiltrating liquid for charged soil particles; v)
Repeated wetting and drying will cause the disintegration of soil clods
through unequal stains and stresses brought about by swelling and
shrinkage and the disruptive effect of escaping air bubbles, however,
unless there is periodically a phase of excessive desiccation leading to
cementation through the dehydration of soil colloids, the fragments
released may not be sufficiently water-stable to result in any long-term
structural benefit; vi)
Prolonged slow desiccation in fine-textured soils will result in lateral
shrinkage and the production of prismatic units. This stage presumably
precedes the development of fragmental cracking within the large
prismatic units; vii)
The prismatic units produced by the progressive drying out of soils in
depth with the production of lateral as well as vertical shrinkage may
create a permanent system of cracks. Dehydration effects localized to
crack faces may stabilize these surfaces. They may be in-filled from the
surface by fine material carried down by gravity or percolating water.
Eventually such cracks provide an effective net-work of drainage
channels but, though plant roots and earthworms may be found within
them, they seem unable to penetrate the surrounding compact, prismatic,
blocks. Thus, prismatic cracking may be said to provide only very
limited access to the subsoil environment.
The Influence of Roots:
Good granulation is often to be associated with a good grass sword but
how far grass roots actually cause fragmentation and how far they merely
exploit its presence is not clear. Clearly grass roots, by incorporating
an abundant supply of fresh organic matter within the soil, could help
to maintain good structure but, only by physically moulding the soil
could they be said to be actually responsible for the creation of the
porous, granular, aggregates with which they are commonly associated. It
seems most probable that roots penetrate soil masses along pre-existing
cracks, which are large enough to be aerobic and thereafter, their
presence ensures a continuous supply of fresh organic residues, which
may contribute to the continued stabilization of exposed structure
faces.
Work with natural and synthetic compounds has shown
the wide range of substances, which can act as particle cements and
thereby might have an influence on aggregate stability. This type of
approach has also produced evidence of the importance of resistance to
decay in the aggregating compound itself. In this many synthetic
polyelectrolytes, such as Krilium have a very real advantage. However,
since the very property which renders them useful — their adhesiveness
— is likely to prevent their penetrating existing structures or mixing
freely with the soil it is clear that unless they are mixed into
aggregates at the time of formation they will be effective only in
stabilizing the surface of existing structures.
GRANULATION PROCESSES
Any process leading to localized pockets of
cementation, when combined with a process involving differential
swelling and shrinkage and the creation of cracks, will undoubtedly
result in fragmentation. This then could be stabilized by the invasion
of plant roots and the further stabilizing effects of products of
organic decay. However, local pockets of cementation are unlikely to
occur in the uniformly moist conditions of fine-textured possibly around
a nucleous of organic matter. What is required is a granulating
mechanism capable of moulding fine particles into immediately stable
aggregates, 2-3 mm. in diameter.
The Influence of the Earthworm:
Though ignored as an agent which might possibly be involved in the
creation of stable soil aggregates the burrowing earthworm would appear
to be the ideal agent for soil granulation, viz: - i)
Burrowing earthworms can invade moist, structureless, fine-textured soil
by ingesting mineral particles; ii)
Ingested mineral matter is cast within the soil to form a coherant nutty
type of aggregate similar to a shellod walnut in size and form. Such
units can be found with the tip of the earthworms tail imprinted as a
perfect countersunk hollow in the surface. Surface casts will rapidly
fragment into cylindrical units 2-3 mm. in diameter; iii)
Earthworms feed on fresh organic residues, which are intimately mixed
with ingested mineral matter and lime secreted from a special gland
within the gut. The normal cast of the feeding earthworm is, therefore,
an ideal size and composition for stable granulation; vi)
Under humid tropical conditions a high level of microbial activity would
lead to a rapid loss of any organic component but solubilized iron and
aluminium would migrate to the surface of aggregates during periods of
drought and there, through dehydration, form a stable coating; v)
Studies of the extent of earthworm casting in British soils suggest
that the amount of soil consumed by the earthworm population under
established grass may generally be of the order of 20 tons/acre and may
rise to 30. This means that, if most of the soil consumed comes from the
top 6 inches, something like 5 % of the topsoil may pass through the gut
of an earthworm each year under established grass. The equivalent figure
for arable would be much less, approx. 1 %. The question is, is this
enough to establish a structural pattern, which can be maintained by
grass roots.
CLIMATE AND STRUCTURE
With drying and wetting, freezing and thawing,
dehydration, clay and organic matter content, soluble iron and aluminium,
level of microbial activity, earthworm activity and surface vegetation
all cited as possible influences on the development of soil structure it
may well be that climate, which has an influence on all these things,
may be correlated with particular manifestations of structural
development; and so it would appear: i)
Cold winters with cracking promoted by freezing; ii)
Cold winters ensuring arrested decay of fresh organic residues; iii)
Short period of intense earthworm activity and microbial decay during a
short warm, moist spring; vi)
Desiccation and arrested decay during the hot, dry and summer; v)
High base status, high organic matter content and adequate clay.
Conditions necessary for the development of wheat is
called"good crumb structure"
i)
Aluminium content of flocculated clay - 10 per cent; ii)
An adequate and fairly continuous supply of fresh organic residues; iii)
A base-status, soil temperature and moisture state conducive to the
presence of burrowing earthworms and a moderate rate of organic decay; vi)
Ideally, deciduous forest or permanent pasture, where cultivations do
not damage what would appear to be a more delicate structure than that
developed in chernozems, and where the surface is continually protected
and there is time for the development of extensive root system.
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